Education

PhD in Environmental, Population and Organismic Biology

University of Colorado

MS in Fisheries Science

National Taiwan University

BS in Aquaculture

National Taiwan University of Marine Science and Technology

Personal Website

https://www.mscd.edu/~biology/facstaff/faculty/liu/liu.html

Published Works

Liu, H., Schroeder, L., Berry, A., Dillon, R. . High levels of mtDNA sequence divergence among isolated populations of Fontigens (Truncatelloidea, Hydrobiidae) in the eastern United States.. (in press). Journal of Molluscan Studies. Liu, H., Hershler, R. (2019). A new species and range extensions for three other species of pebblesnails (Lithoglyphidae, Fluminicola) from the upper Klamath basin, California–Oregon.. Zookeys, 8127-67. . Liu, H., Hershler, R., Hovingh, P. (2018). Molecular evidence enables further resolution of the western North American Pyrgulopsis kolobensis complex (Caenogastropoda: Hydrobiidae).. Journal of Molluscan Studies, 84(1), 103-107. . Johnson, M. J., Schendel, K. R., Ribble, M. B., Liu, H. (2018). Institutional Capacity Building for STEM Teacher Education at an Urban Commuter University . (). Peter Lang--Social Justice Across Contexts in Education series. Hershler, R., Liu, H. (2017). Annotated Checklist of Freshwater Truncatelloidean Gastropods of the Western United States, with an Illustrated Key to the Genera.. (Issue Technical Note 449, pp 142). United States Department of the Interior, Bureau of Land Management National Operations Center. Hershler, R., Liu, H., Forsythe, C., Hovingh, P., Wheeler, K. (2017). Partial revision of the Pyrgulopsis kolobensis complex (Caenogastropoda: Hydrobiidae), with resurrection of P. pinetorum and description of three new species from the Virgin River drainage, Utah.. Journal of Molluscan Studies, 83(2), 161-171. . Hershler, R., Liu, H., Hubbart, N. (2017). Two new species of Fluminicola (Caenogastropoda, Lithoglyphidae) from southwest Oregon, USA, and a range extension for F. multifarious. . Zookeys, 679-20. . Hershler, R., Liu, H., Stevens, L. (2016). A New Springsnail (Hydrobiidae: Pyrgulopsis) from the Lower Colorado River Basin, Northwestern Arizona.. Western North American Naturalist , 76(1), 72-81. . Liu, H., Marceau, D., Hershler, R. (2016). Taxonomic identity of two amnicolid gastropods of conservation concern in lakes of the Pacific Northwest of the USA. . Journal of Molluscan Studies , 8264-471. . Hershler, R., Liu, H., Babbitt, K., Howard, J. (2016). Three new species of western California springsnails previous confused with Pyrgulopsis stearnsiana (Caenogastropoda, Hydrobiidae).. Zookeys, 601-19. . Hershler, R., Liu, H., Simpson, S. J. (2015). Assembly of a micro-hotspot of caenogastropd endemism in the southern Nevada desert, with a description of a new species of Tryonia (Truncatelloidea, Cochliopidae). ZooKeys, 49207-122. http://zookeys.pensoft.net. Hershler, R., Liu, H., Simpson, S. J. (2015). Assembly of a micro-hotspot of caenogastropod endemism in the southern Nevadea desert, with a description of a new species of Tryonia (Cochliopidae). Zookeys, 49207-122. . Liu, H., Hovingh, P., Hershler, R. (2015). Genetic evidence for recent spread of springsnails (Hydrobiidae: Pyrgulopsis) across the Wasatch Divide by stream capture. (in press). Western North American Naturalist. Hershler, R., Liu, H., Carlton, T. J., Cohen, N. A., Davis, B. C., Sorensen, J., Weedman, D. (2015). New discoveries of introduced and cryptogenic fresh and brackish water gastropods (Caenogastropoda: Cochliopidae) in the western United States. Aquatic Invasions, 10(2), 147-156. . Liu, H., Hershler, R., Rossel, C. (2015). Taxonomic status of the Columbia duskysnail (Truncaelloidea, Amnicolidae: Colligyrus). Zookeys, 514-13. . Liu, H., Hershler, R. (2014). Microsatellite primers for a western North American springsnail (Pyrgulopsis robusta): evidence for polyploidy and cross-amplification in P. bruneauensis. Journal of Molluscan Studies/Oxford Press, 80(1), 107-110. . Hershler, R., Landye, J. J., Liu, H., De la Maza-Benignos, M., Carson, W. E. (2014). New species and records of Chihuahuan Desert springsnail, with a new combination for Tryonia brunei. Western North American Naturalist, 74(1), 47-65. . Hershler, R., Liu, H., Howard, J. (2014). Springsnails: a new conservation focus in Western North America. Bioscience, 64(8), 693-700. . Hershler, R., Ratcliffe, V., Liu, H., Lang, B., Hay, M. C. (2014). Taxonomic revision of the Pyrgulopsis gilae (Caenogastropoda, Hydrobiidae) species complex, with descriptions of two new species from the Gila River basin, New Mexico. Zookeys, 4299-85. https://doi.org/10.3897/zookeys.429.7865. Thompson, J., Hershler, R., Liu, H., Simpson, S. J. (2013, Fall). Delineating new species in the Badwater Snail. Society for Advancing Hispanics, Chicanos and Native Americans in Science. Liu, H., Hershler, R., Lang, B., Davis, J. (2013). Molecular evidence for cryptic species in a narrowly endemic western North American springsnail (Pyrgulopsis gilae). Conservation Genetics, 14(4), 917-923. . Wilke, T., Haase, M., Hershler, R., Liu, H., Misof, B., Ponder, W. (2013). Pushing short DNA fragments to the limit: Phylogenetic relationships of 'hydrobioid' gastropods (Caenogastropoda: Rissooidea). Molecular Phylogenetics and Evolution, 66(3), 715-736. . Hershler, R., Liu, H., Bradford, C. (2013). Systematics of a widely distributed western North American springsnail, Pyrgulopsis micrococcus (Caenogastropoda, Hydrobiidae), with descriptions of three new congeners. Zookeys, 3307-52. . Liu, H., Walsh, J., Hershler, R. (2013). Taxonomic clarification and phylogeography of Fluminicola coloradenis Morrison (Caenogastropoda: Lithoglyphidae), a widely ranging western North American pebblesnail. Monographs of the Western North American Naturalist, 67-110. . Hershler, R., Liu, H. (2012). A new species of springsnail (Pyrgulopsis) from the Owyhee River Basin, Nevada. Western North American Naturalist, 72(1), 21-31. . Hershler, R., Liu, H., Clark, W. (2012). Comments on “The absence of genotypic diversity in a successful parthenogenetic invader” by Mark Dybdahl and Devin Drown [Biological Invasions 13 (2011), 1663-1672]. Biological Invasions, 14643-1645. . Liu, H., Marn, J., Hershler, R., Worsfold, T. (2012). Microsatellite evidence for tetraploidy in the New Zealand mudsnail, Potamopyrgus antipodarum (Gray, 1843). Journal of Molluscan Studies/Oxford Press, 78(2), 227-230. . Hershler, R., Liu, H. (2012). Molecular phylogeny of the western North American pebblesnails, genus Fluminicola (Rissooidea: Lithoglyphidae), with description of a new species. Journal of Molluscan Studies/Oxford Press, 78(4), 321-329. . Liu, H., Hershler, R. (2012). Phylogeography of an endangered Western North American springsnail. Conservation Genetics, 13(1), 299-305. . Liu, H. (2011). A federally listed endangered species, Bruneau Hot Springsnail. , Hershler, R., Thompson, G. F., Liu, H. (2011). A large range extension for the monotypic North American aquatic gastropod genus Cincinnatia based on morphologic and molecular evidence. Journal of Molluscan Studies/Oxford University Press, 7732-240. . Hershler, R., Liu, H. (2011). Comments on the paper "Is the track of the Yellowstone hotspot driven by a deep mantle plume? -- Review of volcanism, faulting, and uplift in light of new data" by Kenneth Pierce and Lisa Morgan [J. Volcanol. Geotherm. Res. 188 (2009), 1-25]. Journal of Volcanology and Geothermal Research/Elsevier B.V., 202(3-4), 262-263. . Liu, H. (2011). Cross-species microsatellite amplification. , Hershler, R., Liu, H., Landye, J. (2011). Description of two new genera and four new species of freshwater gastropods (Rissooidea: Cochliopidae) from northeastern Mexico. Journal of Molluscan Studies/Oxford Press, 77-23. . Hershler, R., Liu, H., Landye, J. J. (2011). New species and records of Springsnails (Caenogastropoda: Cochliopidae: Tryonia) from the Chihuahuan Desert (Mexico and United States), in imperiled biodiversity hotspot. Zootaxa/Magnolia Press, 3001-32. . Hershler, R., Liu, H. (2011). Rediscription of Marstonia comalensis (Pilsbry & Ferris, 1906), a poorly known and possibly threatened freshwater gastropod from the Edwards Plateau region (Texas). Zookeys, 77-16. . Hershler, R., Liu, H. (2010). Bruneau Hot Springsnail Gentic Survey. , Hershler, R., Liu, H. (2010). Genetic variation of threatened springsnails from the Gila River basin, New Mexico. , Heshler, R., Liu, H., Clark, H. W. (2010). Microsatellite evidence of invasion and rapid spread of divergent New Zealand mudsnail (Potamopyrgus antipodarum) clones in the Snake River basin, Idaho, USA. Biological Invasions/Springer Science+Business Media, 12(6), 1521-1532. . Hershler, R., Liu, H., Lang, K. B. (2010). Transfer of the Phantom Cave snail, Cochliopa texana, to Pyrgulopsis (Hydrobiidae) and description of a third congener from the Lower Pecos River Basin. Journal of Molluscan Studies/Oxford Press, 76(3), 245-256. . Hershler, R., Liu, H. (2010). Two new, possibly imperiled species of Pyrgulopsis (Gastropoda: Hydrobiidae) from southwestern California. Zootaxa/Magnolia Press, 2343-17. . Liu, H. (2010). Using microsatellite data to detect ploidy level in the New Zealand mudsnail, Potamopyrgus antipodarum (Gray, 1843). , Liu, H., Hershler, R. (2009). Genetic diversity and population structure of the threatened Bliss Rapids snail (Taylorconcha serpenticola). Freshwater Biology/Wiley-Blackwell, 54(6), 1285-1299. . Hershler, R., Liu, H. (2009). New species and records of Pyrgulopsis (Gastropoda: Hydrobiidae) from the Snake River basin, southeastern Oregon: further delineation of a highly imperiled fauna. Zootaxa/Magnolia Press, 2006-22. . Liu, H. (2009). Taxonomic clarification of pebblesnails (Lithoglyphidae: Fluminicola) of the northwestern Great Basin. , Hershler, R., Liu, H., Gustafson, L. D. (2008). A second species of Pyrgulopsis (Hydrobiidae) from the Missouri River Basin, with molecular evidence supporting faunal origin through Pliocene stream capture across the Northern Continental Divide. Journal of Mulloscan Studies/Oxford Press, 74(4), 403-413. . Hershler, R., Liu, H. (2008). Ancient vicariance and recent dispersal of springsnails (Hydrobiidae: Pyrgulopsis) in the Death Valley system, California-Nevada. (439, pp 91-101). Late Cenozoic Drainage History of the Southwestern Great Basin and Lower Colorado River Region: Geologic and Biotic Perspectives/Geological Society of America Special Paper. Liu, H., Hershler, R. (2008). Microsatellite markers for the threatened Bliss Rapids snail (Taylorconcha serpenticola) and cross-amplification in its congener, T. insperata. Molecular Ecology Resources/Wiley-Blackwell, 8(2), 418-420. . Hershler, R., Liu, H. (2008). Phylogenetic relationships of assimineid gastropods of the Death Valley lower Colorado River region: relicts of a late Neogene marine incursion?. Journal of Biogeography/Wiley-Blackwell, 35(10), 1816-1825. . Liu, H., Hershler, R. (2007). A test of the vicariance hypothesis of Western North American freshwater molluscan biogeography. Journal of Biogeography/Wiley-Blackwell, 3434-548. . Hershler, R., Davis, L. C., Kitting, L. C., Liu, H. (2007). Discovery of introduced and cryptogenic cochliopid gastropods in the San Francisco Estuary, California. Journal of Molluscan Studies/Oxford Press, 73(4), 323-332. . Hershler, R., Liu, H., Frest, J. T., Johannes, J. E. (2007). Extensive diversification of pebblesnails (Lithoglyphidae: Fluminicola) in the upper Sacramento River basin, northwestern United States. Zoological Journal of Linnean Society/Wiley-Blackwell, 14971-422. . Hershler, R., Liu, H., Lang, K. B. (2007). Genetic and morphologic variation of the Pecos assiminea, an endangered mollusk of the Rio Grande region, United States and Mexico (Caenogastropoda: Rissoidea: Assimeidae). Hydrobiologia/Springer Science+Business Media, 57917-335. . Ramey, R. R., Wehausen, D. J., Liu, H., Epps, W. C., Carpenter, M. L. (2007). How King et al. (2006) define an "evolutionary distinction" of a mouse subspecies: a response. Molecular Ecology/Wiley-Blackwell, 16(17), 3518-3521. . Hershler, R., Liu, H., Sada, W. D. (2007). Origin and diversification of the soldier meadow springsnails (Hydrobiidae: Pyrgulopsis), a species flock in the northwestern Great Basin, United States. Journal of Molluscan Studies/Oxford Press, 73(2), 167-183. . Hershler, R., Liu, H., Frest, J. T., Johannes, J. E., Clark, H. W. (2006). Genetic structure of the Western North American aquatic gastropod genus Taylorconcha and description of a second species. Journal of Molluscan Studies/Oxford Press, 7267-177. . Wu, K. S., Liu, H. (2006). Kansas Physidae (Gastropoda: Pulmonata: Hygrophila). Malacological Review, 33/349-79. . Ramey, R. R., Wehausen, D. J., Liu, H., Epps, W. C., Carpenter, M. L. (2006). Response to: Should hypotheses testing or selective post hoc interpretation of results guide the allocation of conservation effort?. Animal Conservation/Wiley-Blackwell, 9(3), 244-247. . Ramey, R. R., Liu, H., Epps, W. C., Carpenter, M. L., Wehausen, D. J. (2005). Genetic relatedness of the Preble's meadow jumping mouse (Zapus hudsonius preblei) to nearby subspecies of Z. hudsonius as inferred from variation in cranial morphology, mitochondrial DNA, and microsatellite DNA: implication for taxonomy and conservation. Animal Conservation/Wiley-Blackwell, 8(3), 329-346. . Hershler, R., Mulvey, M., Liu, H. (2005). Genetic variation in the Desert Springsnail (Tryonia porrecta): implications for reproductive mode and dispersal. Molecular Ecology/Wiley-Blackwell, 14755-1766. . Liu, H., Hershler, R. (2005). Molecular systematics and radiation of Western North American Nymphophiline Gastropods. Molecular Phylogenetics and Evolution/Elsevier, 34(2), 284-298. . Hershler, R., Liu, H. (2004). A molecular phylogeny of aquatic gastropods provides a new perspective on biogeographic history of the Snake River region. Molecular Phylogenetics and Evolution/Elsevier, 3227-937. . Hershler, R., Liu, H. (2004). Taxonomic reappraisal of species assigned to the North American freshwater Gastropod Subgenus Natricola Gregg & Taylor (Rissooidea: Hydrobiidae). The Veliger/California Malacozoological Society, 47(1), 66-81. . Liu, H., Hershler, R., Clift, K. (2003). Mitochondrial DNA sequences reveal extensive cryptic diversity within a western American springsnail. Molecular Ecology/Wiley-Blackwell, 12771-2782. . Hershler, R., Liu, H., Thompson, G. F. (2003). Phylogenetic relationships of North American nymphophiline gastropods based on mitochondrial DNA sequences. Zoologica Scripta/Blackwell Publishing, 3257-366. . Hershler, R., Frest, J. T., Liu, H., Johannes, J. E. (2003). Rissooidean snails from the Pit River Basin, California. The Veliger/California Malacozooligical Society, 46(4), 275-304. . Hershler, R., Liu, H., Landye, J. J. (2002). A new species of aquatic snail of the genus Ermopyrgus (Hydrobiidae: Cochliopinae) from the Chihuahuan Desert, Mexico: Phylogenetic relationships and biogeography. The Journal of Molluscan Studies/Oxford Press, 68-15. . Hershler, R., Liu, H., Stockwell, A. C. (2002). Juturnia, a new genus of aquatic gastropods (Rissooidea: Hydrobiidae) from the North American Southwest: Phylogenetic relationships and biogeography. Proceedings of the Biological Society of Washington/Allen Press, 115(1), 171-188. . Kandl, L. K., Liu, H., Butler, S. R., Hoeh, R. W., Mulvey, M. (2001). A genetic approach to resolving taxonomic ambiguity: Pleurobema (Bivalvia: Unionidae) of the eastern Gulf Coast. Malacologia/Institute of Malacology, 43(1-2), 87-101. . Liu, H., Hershler, R., Thompson, G. F. (2001). Phylogenetic relationships of the Cochliopinae (Rissooidea: Hydrobiidae): an enigmatic group of Aquatic Gastropods. Molecular Phylogenetic and Evolution/Elsevier, 21(1), 17-25. . Hershler, R., Mulvey, M., Liu, H. (1999). Biogeography in the Death Valley region: Evidence from springsnails (Hydrobiidae: Tryonia). Zoological Journal of Linnean Society/Wiley-Blackwell, 12635-354. . Hershler, R., Liu, H., Mulvey, M. (1999). Phylogenetic relationships within the aquatic snail genus Tryonia: Implications for biogeography of the North American Southwest. Molecular Phylogenetics and Evolution/Elsevier, 13(2), 377-391. . Mulvey, M., Liu, H., Kandl, L. K. (1998). Application of molecular genetic markers to conservation of freshwater mussels. Journal of Shellfish Research/The National Shellfisheries Association, 17(5), 1395-1405. . Aars, J., Ims, A. R., Liu, H., Mulvey, M., Smith, M. (1998). Bank voles in linear habitats shows restricted gene flow as revealed by mtDNA sequences. Molecular Ecology/Wiley-Blackwell, 7383-1389. . Liu, H., Mitton, B. J., Herrmann, J. S. (1996). Genetic differentiation and management recommendations for the freshwater mussel, Pyganodon grandis Say 1829. American Malacological Bulletin/American Malacological Society, 13(1/2), 117-125. . Liu, H., Mitton, B. J., Wu, K. S. (1996). Paternal mitochondrial DNA differentiation far exceeds maternal mitochondrial DNA and allozyme differentiation in the freshwater mussel, Anodonta grandis grandis. Evolution/Wiley-Blackwell, 50(2), 952-957. . Liu, H., Mitton, B. J. (1996). Tissue-specific maternal and paternal mitochondrial DNA in the freshwater mussel, Anodonta grandis grandis Say 1829. The Journal of Molluscan Studies/Oxford Press, 6293-394. . Liu, H., Liao, Y. K., Wu, L. W. (1993). Comparison of the reproductive ecology of Meretrix lusoria from Tam-Shui and Lu-Kang area. Bulletin of Malacology/Malacological Society of Republic of China, 179-77. . Liu, H. (1993). Diagnostic genetic loci for species in the genus Physella. Malacological Review/University of Michigan, 26-8. . Wu, L. W., Liu, H., Liao, Y. K. (1993). Morphological and anatomical studies of Meretrix lusoria from Taiwan (Bivalvia: Veneridae). Bulletin of Malacology/Malacological Society of Republic of China, 171-98. . Wu, L. W., Liu, H. (1993). Studies on the gonad development of Meretrix lusoria from Taiwan (Bivalvia: Veneridae). Bulletin of Malacology/Malacological Society of Republic of China, 179-86. . Liu, H., Mitton, B. J. (1993). Techniques to reveal restriction fragment length polymorphisms (RFLPs) in the genus Physa. Malacological Review/University of Michigan, 269-90. . Wu, L. W., Liu, H. (1989). Malacological research on Meretrix resources in Taiwan. II. History review and evaluation on the studies of the Taiwan Meretrix. Bulletin of Malacology/Malacological Society of Republic of China, 149-61. . Wu, L. W., Liu, H. (1988). Malacological research on Meretrix resources in Taiwan. I. Preliminary report on isozyme study. Bulletin of Malacology/Malacological Society of Republic of China, 13-7. .